Srygley, R.B., Dudley, R., Hernandez, E.J., Kainz, F., Riveros, A.J. and C.P. Ellington. (2023). Quantifying the aerodynamic power required for flight and testing for adaptive wind drift in passion-vine butterflies Heliconius sara (Lepidoptera: Nymphalidae). Insects 14:112. (12 pp.) [link]


Brown, C.E., Sathe, E.A., Dudley, R. and S.M. Deban. (2022a). Aerial maneuvering by plethodontid salamanders spanning an arboreality gradient. Journal of Experimental Biology 225, jeb244598. [link]

Brown, C., Sathe, E.A., Dudley, R. and S.M. Deban. (2022b). Gliding and parachuting by arboreal salamanders. Current Biology 32: R453-454. [link]

Campbell, C., Maro, A., Weaver, V. and R. Dudley. (2022). Dietary ethanol ingestion by free-ranging spider monkeys (Ateles geoffroyi). Royal Society Open Science 9:211729. [link]

Kou, G., Wang, Y., Dudley, R., Wu, Y. and D. Li. (2022). Coping with captivity: takeoff speed and load-lifting capacity are unaffected by substantial changes in body condition for a passerine bird. Journal of Experimental Biology 225, jeb244642 (7 pp.) [link]

Louis, L.D., Bowie, R.C.K. and R. Dudley. (2022). Wing and leg bone microstructure reflects migratory demands in resident and migrant populations of the Dark-eyed Junco (Junco hyemalis). Ibis 164:132-150. [link]

Maro, A. and R. Dudley. (2022). Non-random distribution of ungulate salt licks relative to distance from North American oceanic margins. Journal of Biogeography 49:254-260. [link]


Carn, D., Lanaspa, M.A., Benner, S.A., Andrews, P., Dudley, R., Andres-Hernando, A., Tolan, D.R and R.J. Johnson. (2021). The role of thrifty genes in the origin of alcoholism: a narrative review and hypothesis. Alcoholism: Clinical and Experimental Research 45:1519–1526. [link]

Dudley, R. (2021). A morphofunctional hypothesis for selection on EDAR V370A and associated elements of sinodonty. Dental Anthropology 34:49-54. [link]

Dudley, R. and A. Maro. (2021). Human evolution and dietary ethanol. Nutrients 13:2419 (9 pp.) [link]


Dudley, R. (2020). The natural biology of dietary ethanol, and its implications for primate evolution. In: Alcohol and Humans: A Long and Social Affair (eds. K. Hockings and R. Dunbar), pp. 10–23. Oxford: Oxford University Press. [link]

Ortega-Jimenez, V. M., Herbst, E.C., Leung, M. and R. Dudley. (2020). Natural barriers: waterfall transit by small flying animals. Royal Society Open Science 7:201185 (9 pp.) [link]

Su, G., Dudley, R., Pan, T., Zheng, M., Peng, L. and Q. Li. (2020). Maximum aerodynamic force production by the wandering glider dragonfly (Pantala flavescens, Libellulidae). Journal of Experimental Biology 223:218552 (8 pp.) [link]

Zeng, Y., Chang, S.W., Williams, J.Y., Nguyen, L.Y.-N., Tang, J., Naing, G., Kazi, C. and R. Dudley. (2020). Canopy parkour: movement ecology of post-hatch dispersal in a gliding nymphal stick insect (Extatosoma tiaratum). Journal of Experimental Biology 223:226266 (12 pp.) [link]

Zeng, Y., O’Malley, C., Singhal, S., Rahim, F., Park, S., Chen, X. and R. Dudley. (2020). A tale of winglets: evolution of flight morphology in stick insects. Frontiers in Ecology and Evolution 8:121 (15 pp.) [link]


Badger, M., Wang, H. and R. Dudley. (2019). Avoiding topsy-turvy: how Anna’s Hummingbirds (Calypte anna) fly through upward gusts. Journal of Experimental Biology 222:176263, 10 pp. [link]

Dudley, R. and R.J. Wootton. (2019). Charles Porter Ellington (1952-2019). Journal of Experimental Biology 222:218628 (2 pp.) [link]

Ortega-Jimenez, V.M. and R. Dudley. (2019). Superb autorotator: rapid decelerations in impulsively launched samaras. Journal of the Royal Society Interface 16:20181456, 5 pp. [link]

Rico-Guevara, A., Rubega, M.A., Hurme, K.J. and R. Dudley. (2019). Shifting paradigms in the mechanics of nectar feeding and hummingbird bill morphology. Integrative Organismal Biology 10.1093/iob/oby006, 15 pp. [link] 

Rico-Guevara, A., Sustaita, D., Gussekloo, S., Olsen, A., Bright, J., Corbin, C. and R. Dudley. (2019). Feeding in birds: thriving in terrestrial, aquatic, and aerial niches. In: Feeding in Vertebrates (ed. V.L. Bels and I.Q. Whishaw), pp. 643-693. Berlin: Springer Verlag. [link]

Wang, Y., Yin, Y., Ge, S., Li, M., Zhang, Q., Li, J., Wu, Y., Li, D. and R. Dudley. (2019). Limits to load-lifting performance in a passerine bird: the effects of intraspecific variation in morphological and kinematic parameters. PeerJ 7:e8048, 12 pp. [link]


Dudley, R. and G. Pass. (2018). Wings and powered flight: core novelties in insect evolution. Arthropod Structure & Development 47:319-321. [link]

Ortega-Jimenez, V.M. and R. Dudley. (2018). Ascending flight and decelerating vertical glides in Anna’s Hummingbirds. Journal of Experimental            Biology 14:20180152. [link]

Skowron Volponi, M.A.A., McLean, D.J., Volponi, P. and R. Dudley. (2018). Moving like a model: mimicry of hymenopteran flight trajectories by clearwing moths of Southeast Asian rainforests. Biology Letters 14:20180152. [link]

Yanoviak, S.P. and R. Dudley. (2018). Jumping and the aerial behavior of aquatic mayfly larvae (Myobaetis ellenae, Baetidae). Arthropod Structure & Development 47:370-374. [link]


Ortega-Jimenez, V.M., von Rabenau, L. and R. Dudley. (2017). Escape jumping by three age classes of water striders from smooth, wavy and bubbling water surfaces. Journal of Experimental Biology 220:2809-2815. [link]

Ortega-Jimenez, V.M., Martín-Alcántara, A., Fernandez-Feria, R. and R. Dudley. (2017). On the autorotation of animal wings. Journal of the Royal Society Interface 14:20160870. (10 pp.) [link]

Anderson, P., Berridge, V., Conrod, P., Dudley, R., Hellman, M., Lachenmeier, D., Lingford-Hughes. A., Miller. D., Rehm, J., Room, R., Schmidt, L., Sullivan, R., Ysa, T. and A. Gual. (2017). Reframing the science and policy of nicotine, illegal drugs and alcohol – conclusions of the ALICE RAP Project. F1000Research 6:289. (13 pp.) [link]

Muijires, F.T., Chang, S., van Veen, W., Spitzen, J., Biemans, B.T., Koehl, M.A.R. and R. Dudley. (2017). Escaping blood-fed malaria mosquitoes minimize tactile detection without compromising on take-off speed. Journal of Experimental Biology 220:3751-3762. [link]

Skandalis, D.A., Bahlman, J.W., Groom, D., Segre, P.S., Welch, K.C., Witt, C.C., McGuire, J.A., Dudley, R., Lentink, D. and D.L. Altshuler. (2017). The biomechanical origin of extreme wing allometry in hummingbirds. Nature Communications 8:1047 (9 pp.) [link]

Zeng, Y., Lam, K., Chen, Y., Gong, M., Xu, Z. and R. Dudley. (2017). Biomechanics of aerial righting in wingless nymphal stick insects. Interface Focus 7:20160075 (12 pp.) [link]


Badger, M., Ortega-Jimenez, V., Wang, H. and R. Dudley. (2016). Into rude air: hummingbird flight performance in variable aerial environments. Philosophical Transactions of the Royal Society of London B 371:20150387 (7 pp.) [link]

Ortega-Jimenez, V.M., Arriaga-Ramírez, S. and R. Dudley. (2016). Meniscus ascent by thrips (Thysanoptera). Biology Letters 12:20160279 (4 pp.) [link]

Sun, Y.-F., Ren, Z.-P., Wu, Y.-F., Lei, F.-M., Dudley, R. and D.-M. Li. (2016). Flying high: limits to flight performance by sparrows on the Qinghai-Tibetan plateau. Journal of Experimental Biology 219:3642-3648. [link]


Badger, M., Ortega-Jimenez, V.M., von Rabenau, L., Smiley, A. and R. Dudley. (2015). Electrostatic charge on flying hummingbirds and its potential role in pollination. PLOS ONE doi:10.1371/journal.pone.0138003 (11 pp.) [link]

Full, R.J., Dudley, R., Koehl, M.A.R., Libby, T. and C. Schwab. (2015). Interdisciplinary laboratory course facilitating knowledge integration, mutualistic teaming, and original discovery. Integrative and Comparative Biology 5:912-925. [link]

Munk, Y., Yanoviak, S.P., Koehl, M.A.R. and R. Dudley. (2015). The descent of ant: field-measured three-dimensional trajectories of gliding ants.      Journal of Experimental Biology 218:1393-1401. doi:10.1242/jeb.106914 [link]

Yanoviak, S.P., Munk, Y. and R. Dudley. (2015). Arachnid aloft: directed aerial descent in a Neotropical canopy spider. Journal of the Royal Society Interface 12:20150534 (5 pp.) doi:10.1098/rsif.2015.0534 [link]

Zeng, Y., Lin, Y., Abundo, A. and R. Dudley. (2015). The visual ecology of directed aerial descent in first-instar nymphs of the stick insect Extatosoma tiaratum. Journal of Experimental Biology 218:2305-2314. doi:10.1242/jeb.109553 [link]


Dillon, M.E. and R. Dudley. (2014). Surpassing Mt. Everest: extreme flight performance of alpine bumble-bees. Biology Letters doi:10.1098/rsbl.2013.0922 (4 pp.) [link]

Dudley, R. (2014). The Drunken Monkey: Why We Drink and Abuse Alcohol. Berkeley: University of California Press. 154 pp. [link]

Evangelista, D., Cam, S., Huynh, T., Krivitskiy, I. and R. Dudley. (2014). Ontogeny of aerial righting and wing flapping in juvenile birds. Biology Letters doi:10.1098/rsbl.2014.0497 (4 pp.) [link]

Evangelista, D., Cam, S., Huynh, T., Kwong, A., Mehrabani, H., Tse, K. and R. Dudley. (2014). Shifts in stability and control effectiveness during evolution of Paraves support aerial maneuvering hypotheses for flight origins. PeerJ 2:e632 (25 pp.) [link]

Kim, E.J., Wolf, M., Ortega-Jimenez, V.M. and R. Dudley. (2014). Hovering performance of Anna’s Hummingbirds (Calypte anna) in ground effect. Journal of the Royal Society Interface 11: 20140505 (8 pp.) [link]

McGuire, J.A., Witt, C.C., Van Remsen, J., Corl, A., Rabosky, D.L., Altshuler, D.L. and R. Dudley. (2014). Molecular phylogenetics and the diversification of hummingbirds (Apodiformes: Trochilidae). Current Biology doi:10.1016/j.cub.2014.03.016 (7 pp.) [link]

Ortega-Jimenez, V.M., Wolf, M., Variano, E.A. and R. Dudley. (2014). Into turbulent air: size-dependent effects of von Kármán vortex streets on hummingbird flight kinematics and energetics. Proceedings of the Royal Society B doi:10.1098/rspb.2014.0180 (10 pp.) [link]

Srygley, R.B., Dudley, R., Oliveira, E.G. and A.J. Riveros. (2014). El Niño, host plant growth, and migratory butterfly abundance in a changing climate. Biotropica 46:90-97. [link]

Xu, X., Zhou, Z., Dudley, R., Mackem, S., Chuong, C.-M., Erickson, G.M. and D.J. Varricchio. (2014). An integrative approach to understanding bird origins. Science 346:1253293 doi:10.1126/science.1253293 (12 pp.) [link]


Ortega-Jimenez, V.M. and R. Dudley. (2013). Rapid spiderweb deformation induced by electrostatically charged insects. Scientific Reports doi:10.1038/srep02108 (4 pp.) [link]

Projecto-Garcia, J., Natarajan, C., Moriyama, H., Weber, R.E., Fago, A., Cheviron, Z.A., Dudley, R., McGuire, J.A., Witt, C.C. and J.F. Storz. (2013). Repeated elevational transitions in hemoglobin function during the evolution of Andean hummingbirds. Proceedings of the National Academy of Sciences USA 110:20669-20674. [link]

Sapir, N. and R. Dudley. (2013). Implications of floral orientation for flight kinematics and metabolic expenditure of hover-feeding hummingbirds. Functional Ecology 27:227-235. [link]

Wolf, M., Ortega-Jimenez, V.M. and R. Dudley. (2013). Structure of the vortex wake in hovering Anna’s Hummingbirds (Calypte anna). Proceedings of the Royal Society B doi:10.1098/rspb.2013.2391 (7 pp.) [link]


Dudley, R., Kaspari, M. and S.P. Yanoviak. (2012). Lust for salt in the western Amazon. Biotropica 44:6-9. [link]

Medeiros, M.J. and R. Dudley. (2012). Jumping performance in flightless Hawaiian grasshopper moths (Xyloryctidae: Thyrocopa spp.). Proceedings of the Hawaiian Entomological Society 44:55-61. [link]

Ortega-Jimenez, V.M. and R. Dudley. (2012a). Aerial shaking performance of wet Anna’s Hummingbirds. Journal of the Royal Society Interface 9:1093-1099. [link]

Ortega-Jimenez, V.M. and R. Dudley. (2012b). Flying in the rain: hovering performance of Anna’s Hummingbirds under varied precipitation. Proceedings of the Royal Society B 279:3996-4002. [link]

Sapir, N. and R. Dudley. (2012). Backward flight in hummingbirds employs unique kinematic adjustments and entails low metabolic cost. Journal of Experimental Biology 215:3603-3611. [link]


Buermann, W., Chaves, J.A., Dudley, R., McGuire, J.A., Smith, T.B. and D.L. Altshuler. (2011). Projected changes in elevational distribution and flight performance of montane Neotropical hummingbirds in response to climate change. Global Change Biology 17:1671-1680. [link]

Dudley, R. and S.P. Yanoviak. (2011). Animal aloft: the origins of aerial behavior and flight. Integrative and Comparative Biology 51:926-936. [link]

Fernández, M.J., Dudley, R. and F. Bozinovic. (2011). Comparative energetics of the Giant Hummingbird (Patagona gigas). Physiological and Biochemical Zoology 84:333-340. [link]

Jusufi, A., Zeng, Y., Full, R.J. and R. Dudley. (2011). Aerial righting reflexes in flightless animals. Integrative and Comparative Biology 51:937-943. [link]

McGuire, J.A. and R. Dudley. (2011). The biology of gliding in flying lizards (genus Draco) and their fossil and extant analogs. Integrative and Comparative Biology 51:983-990. [link]

Peterson, K., Birkmeyer, P., Dudley, R. and R. Fearing. (2011). A wing-assisted running robot and implications for avian flight evolution. Bioinspiration & Biomimetics 6 046008 (8 pp.) [link]

Yanoviak, S.P., Munk, Y. and R. Dudley. (2011). Evolution and ecology of directed aerial descent in arboreal ants. Integrative and Comparative Biology 51:944-956. [link]


Altshuler, D.L., Dudley, R., Heredia, S.M. and J.A. McGuire. (2010). Allometry of hummingbird lifting performance. Journal of Experimental Biology 213:725-734. [link]

Buchwald, R. and R. Dudley. (2010). Limits to vertical force and power production in bumblebees (Hymenoptera: Bombus impatiens). Journal of Experimental Biology 213:426-432. [link]

Clark, C.J. and R. Dudley. (2010). Hovering and forward flight energetics in Anna’s and Allen’s Hummingbirds. Physiological and Biochemical Zoology 83:654-662. [link]

Evangelista, D., Fernández, M.J., Berns, M.S., Hoover, A. and R. Dudley. (2010). Hovering energetics and thermal balance in Anna’s Hummingbirds (Calypte anna). Physiological and Biochemical Zoology 83:406-413. [link]

Sane, S.P., Srygley, R.B. and R. Dudley. (2010). Antennal regulation of migratory flight in the Neotropical moth Urania fulgens. Biology Letters 6:406-409. [link]

Srygley, R.B., Dudley, R., Oliveira, E.G., Aizprúa, R., Pelaez, N.Z. and A.J. Riveros. (2010). El Niño and dry season rainfall influence hostplant phenology and an annual butterfly migration from Neotropical wet to dry forests. Global Change Biology 16:936-945. [link]

Yanoviak, S.P., Munk, Y., Kaspari, M. and R. Dudley. (2010). Aerial manoeuvrability in wingless gliding ants (Cephalotes atratus). Proceedings of the Royal Society B 277:2199-2204. [link]


Bradley, T.J., Briscoe, A.D., Brady, S., Contreras, H.L., Danforth, B.N., Dudley, R., Grimaldi, D., Harrison, J.F., Kaiser, A., Merlin, C., Reppert, S.M., VandenBrooks, J.M. and S.P. Yanoviak. (2009). Episodes in insect evolution. Integrative and Comparative Biology 49:590-606. [link]

Clark, C.J. and R. Dudley. (2009). Flight costs of long, sexually selected tails in hummingbirds. Proceedings of the Royal Society B 276:2109-2115. [link]

Combes, S.A. and R. Dudley. (2009). Turbulence-driven instabilities limit insect flight performance. Proceedings of the National Academy of Sciences USA 106:9105-9108. [link]

Dickinson, M.H. and R. Dudley. (2009). Flight. In: Encyclopedia of Insects, 2nd ed. (eds. V. Resh and R. Cardé), pp. 364-372. San Diego: Academic Press.

Kaspari, M., Yanoviak, S.P., Dudley, R., Yuan, M. and N. Clay. (2009). Sodium shortage as a constraint on the carbon cycle in an inland tropical rainforest. Proceedings of the National Academy of Sciences USA 106:19405-19409. [link]

McGuire, J.A., Witt, C.C., Remsen, J.V., Dudley, R. and D.L. Altshuler. (2009). A higher-level taxonomy for hummingbirds. Journal of Ornithology 150:155-165. [link]

Yanoviak, S.P., Kaspari, M. and R. Dudley. (2009). Gliding hexapods and the origins of insect aerial behaviour. Biology Letters 5:510-512. [link]


Dudley, R. and R.B. Srygley. (2008). Airspeed adjustment and lipid reserves in migratory Neotropical butterflies. Functional Ecology 22:264-270. [link]

Kaspari, M., Yanoviak, S.P. and R. Dudley. (2008). On the biogeography of salt limitation: a study of ant communities. Proceedings of the National Academy of Sciences USA 105:17848-17851. [link]

Kunz, T.H., Gauthreaux, S.A., Hristov, N.I., Horn, J.W., Jones, G., Kalko, E.K.V., Kelly, T.A., Larkin, R.P., McCracken, G.F., Swartz, S.M., Srygley, R.B., Dudley, R., Westbrook, J.K. and M. Wikelski. (2008). Aeroecology: probing and modelling the aerosphere. Integrative and Comparative Biology 48:1-11. [link]

Mazeh, S., Korine, C., Pinshow, B. and R. Dudley. (2008). Does ethanol in fruit influence feeding in the frugivorous yellow-vented bulbul (Pycnonotus xanthopygos)? Behavioural Processes 77:369-375. [link]

Srygley, R.B. and R. Dudley. (2008). Optimal strategies for insects migrating in the flight boundary layer: mechanisms and consequences. Integrative and Comparative Biology 48:119-133. [link]

Yanoviak, S.P., Kaspari, M., Dudley, R. and G. Poinar. (2008). Parasite-induced fruit mimicry in a tropical canopy ant. American Naturalist 171:536-544. [link]


Dudley, R. (2007). Air. In: Encyclopedia of Tidepools and Rocky Shores (eds. M.W. Denny and S.D. Gaines), pp. 8 – 10. Berkeley: University of California Press.

Dudley, R., Byrnes, G., Yanoviak, S.P., Borrell, B.J., Brown, R. and J.A. McGuire. (2007). Gliding and the functional origins of flight: biomechanical novelty or necessity? Annual Review of Ecology, Evolution, and Systematics 38:179-201. [link]


Altshuler, D.L. and R. Dudley. (2006a). Adaptations to life at high elevation: an introduction to the symposium. Integrative and Comparative Biology 46:3-4. [link]

Altshuler, D.L. and R. Dudley. (2006b). The physiology and biomechanics of avian flight at high altitude. Integrative and Comparative Biology 46:62-71. [link]

Srygley, R.B., Dudley, R., Oliveira, E.G. and A. Riveros. (2006). Experimental evidence for a magnetic sense in Neotropical migrating butterflies (Lepidoptera: Pieridae). Animal Behaviour 71:183-191. [link]

Sánchez, F., Korine, C., Steeghs, M., Laarhoven, L.-J., Harren, F.J.M., Cristescu, S.M., Dudley, R. and B. Pinshow. (2006). Ethanol and methanol as possible odor cues for Egyptian fruit bats (Rousettus aegyptiacus). Journal of Chemical Ecology 32:1289-1300. [link]

Dudley, R., Huey, R.B. and D.R. Carrier. (2006). Living History of Physiology: Carl Gans. Advances in Physiology Education 30:102-107. [link]

Dillon, M., Frazier, M.R. and R. Dudley. (2006). Into thin air: physiology and evolution of alpine insects. Integrative and Comparative Biology 46:49-61. [link]

Yanoviak, S.P. and R. Dudley. (2006). The role of visual cues in directed aerial descent of Cephalotes atratus workers (Hymenoptera: Formicidae). Journal of Experimental Biology 209:1777-1783. [link]


Dudley, R. (2005). Evolutionary and historical aspects of ethanol ingestion. In: Comprehensive Handbook of Alcohol-Related Pathology, Volume 1. General aspects of alcohol and mechanisms of disease (eds. V.R. Preedy and R.R. Watson), pp. 3-13. London: Academic Press.

McGuire, J.A. and R. Dudley. (2005). The cost of living large: comparative gliding performance in flying lizards (Agamidae: Draco). American Naturalist 166:93-106. [link]

Borrell, B.J., Goldbogen, J. and R. Dudley. (2005). Flapping aquatic propulsion at low Reynolds numbers: swimming kinematics of the Antarctic pteropod, Clione antarctica. Journal of Experimental Biology 208:2939-2949. [link]

Borrell, B.J., LaDuc, T.J. and R. Dudley. (2005). Respiratory cooling in rattlesnakes. Comparative Biochemistry and Physiology A 140:471-476. [link]

Stiles, F.G., Altshuler, D.L. and R. Dudley. (2005). On the wing morphology and flight behavior of some North American hummingbird species. The Auk 122:872-886. [link]

Yanoviak, S.P., Dudley, R. and M. Kaspari. (2005). Directed aerial descent in canopy ants. Nature 433:624-626. [link]


Altshuler, D.L., Dudley, R. and J.A. McGuire. (2004). Resolution of a paradox: hummingbird flight at high elevation does not come without a cost. Proceedings of the National Academy of Sciences USA 101:17731-17736. [link]

Altshuler, D.L., Stiles, F.G. and R. Dudley. (2004). Of hummingbirds and helicopters: hovering costs, competitive ability and foraging strategies. American Naturalist 163:16-25. [link]

Dudley, R. (2004). Ethanol, fruit ripening, and the historical origins of human alcoholism in primate frugivory. Integrative and Comparative Biology 44:315-323. [link]

Dudley, R. and M.H. Dickinson. (2004). The comparative biology of ethanol consumption: an introduction to the symposium. Integrative and Comparative Biology 44:267-268. [link]

Altshuler, D.L., Dudley, R. and C.P. Ellington. (2004). Aerodynamic forces of revolving hummingbird wings and wing models. Journal of Zoology, London 264:327-332. [link]

Childress, S. and R. Dudley. (2004). Transition from ciliary to flapping mode in a swimming mollusc: flapping flight as a bifurcation in Reω. Journal of Fluid Mechanics 498:257-288. [link]

Dillon, M.E. and R. Dudley. (2004). Allometry of maximum vertical force production during hovering flight of Neotropical orchid bees (Apidae: Euglossini). Journal of Experimental Biology 207:417-425. [link]

Roberts, S.P., Harrison, J.F. and R. Dudley. (2004). Allometry of kinematics and energetics in Carpenter Bees (Xylocopa varipuncta) hovering in variable-density gases. Journal of Experimental Biology 207:993-1004. [link]

Sánchez, F., Korine, C., Pinshow, B. and R. Dudley. (2004). The possible roles of ethanol in the relationship between plants and frugivores: first experiments with Egyptian fruit bats. Integrative and Comparative Biology 44:290-294. [link]

Stephens, D. and R. Dudley. (2004). The drunken monkey hypothesis. Natural History 113(10):40-44.


Altshuler, D.L. and R. Dudley. (2003). Kinematics of hummingbird hovering flight along simulated and natural elevational gradients. Journal of Experimental Biology 206:3139-3147. [link]

Berner, R.A., Beerling, D.J., Dudley, R., Robinson, J.M. and R.A. Wildman. (2003). Phanerozoic atmospheric oxygen. Annual Review of Earth and Planetary Sciences 31:105-134. [link]

Dickinson, M.H. and R. Dudley. (2003). Flight. In: Encyclopedia of Insects (eds. V. Resh and R. Cardé), pp. 416-426. San Diego: Academic Press.


Altshuler, D.L. and R. Dudley. (2002). The ecological and evolutionary interface of hummingbird flight physiology. Journal of Experimental Biology 205:2325-2336. [link]

Dudley, R. (2002a). Fermenting fruit and the historical ecology of ethanol ingestion: is alcoholism in modern humans an evolutionary hangover? Addiction 97:381-388. [link]

Dudley, R. (2002b). Mechanisms and implications of animal flight maneuverability. Integrative and Comparative Biology 42:135-140. [link]

Dudley, R. and Y. Winter. (2002). Hovering flight mechanics of Neotropical flower bats (Phyllostomidae: Glossophaginae) in normo- and hypodense gas mixtures. Journal of Experimental Biology 205:3669-3677. [link]

Dudley, R., Srygley, R.B., Oliveira, E.G. and P.J. DeVries. (2002). Flight speeds, lipid reserves, and predation of the migratory moth Urania fulgens (Uraniidae). Biotropica 34:120-126. [link]

Moon, B.R., LaDuc, T.J., Dudley, R. and Chang, A. (2002). A twist to the rattlesnake tail. In: Topics in Functional and Ecological Vertebrate Morphology (eds. P. Aerts, R. Van Damme, K. D’Août and A. Herrel), pp. 63-76. Shaker Publishing, Maastricht.


Dudley, R. (2001a). Limits to human locomotor performance: phylogenetic origins and comparative perspectives. Journal of Experimental Biology 204:3235-3240. [link]

Dudley, R. (2001b). The biomechanics and functional diversity of flight. In: Insect Movement: Mechanisms and Consequences (ed. I. Woiwod and D. Reynolds), pp. 19-41. Wallingford: CAB International.


Dudley, R. (2000a). Evolutionary origins of human alcoholism in primate frugivory. Quarterly Review of Biology 75:3-15. [link]

Dudley, R. (2000b). The Biomechanics of Insect Flight: Form, Function, Evolution. Princeton: Princeton University Press. 476 pp. [link]

Dudley, R. (2000c). The evolutionary physiology of animal flight: paleobiological and present perspectives. Annual Review of Physiology 62:135-155. [link]

Vermeij, G.J. and R. Dudley. (2000). Why are there so few evolutionary transitions between aquatic and terrestrial ecosystems? Biological Journal of the Linnean Society 70:541–554. [link]


Chai, P. and R. Dudley. (1999). Maximum flight performance of hummingbirds: capacities, constraints, and trade-offs. American Naturalist 153:398-411. [link]

Dudley, R. (1999). Unsteady aerodynamics. [Perspectives: Biomechanics]. Science 284:1937-1938. [link]

Gans, C., Dudley, R., Aguilar, N.M. and J.B. Graham. (1999a). Late Paleozoic atmospheres and biotic evolution. Historical Biology 13:199-219. [link]

Gans, C., Dudley, R., Aguilar, N.M. and J.B. Graham. (1999b). The pre-Devonian carbon dioxide crash, the late Paleozoic oxygen pulse, and associated shifts in ventilatory mechanisms. In: Water/Air Transition in Biology (ed. A.K. Mittal, F.B. Eddy, and J.S. Datta Munshi), pp. 31-43. New Delhi: Oxford & IBH Publishing Co. [link]


Chai, P., Chang, A.C. and R. Dudley. (1998). Flight thermogenesis and energy conservation in hovering hummingbirds. Journal of Experimental Biology 201:963-968. [link]

Dudley, R. (1998). Atmospheric oxygen, giant Paleozoic insects and the evolution of aerial locomotor performance. Journal of Experimental Biology 201:1043-1050. [link]

Oliveira, E.G., Srygley, R.B. and R. Dudley. (1998). Do neotropical migrant butterflies navigate using a solar compass? Journal of Experimental Biology 201:3317-3331. [link]


Chai, P., Chen, J.S.C. and R. Dudley. (1997). Transient hovering performance of hummingbirds under conditions of maximal loading. Journal of Experimental Biology 200:921-929. [link]

Graham, J.B., Aguilar, N., Dudley, R. and C. Gans. (1997). The late Paleozoic atmosphere and the evolutionary physiology of tetrapods. In: Amniote Origins: Completing the Transition to Land (eds. S.S. Sumida and K.L.M. Martin), pp. 141-167. New York: Academic Press.

Jaramillo, C., Rand, A.S., Ibáñez, R. and R. Dudley. (1997). Elastic structures in the vocalization apparatus of the túngara frog, Physalaemus pustulosus (Leptodactylidae). Journal of Morphology 233:287-295. [link]


Chai, P. and R. Dudley. (1996). Limits to flight energetics of hummingbirds hovering in hypodense and hypoxic gas mixtures. Journal of Experimental Biology 199:2285-2295.[link]

Chai, P., Harrykissoon, R. and R. Dudley. (1996). Hummingbird hovering performance in hyperoxic heliox: effects of body mass and sex. Journal of Experimental Biology 199:2745–2755. [link]

Dudley, R. and G.H. Adler. (1996). Biogeography of milkweed butterflies (Nymphalidae: Danainae) and mimetic patterns on tropical Pacific archipelagos. Biological Journal of the Linnean Society 57:317-326. [link]

Dudley, R. and P. Chai. (1996). Animal flight mechanics in physically variable gas mixtures. Journal of Experimental Biology 199:1881-1885. [link]

Grodnitsky, D.L. and R. Dudley. (1996). Vortex visualization during free flight of heliconiine butterflies (Lepidoptera: Nymphalidae). Journal of the Kansas Entomological Society 69:199-203. [link]

Srygley, R.B., Oliveira, E. and R. Dudley. (1996). Wind drift compensation, flyways, and conservation of diurnal, migrant Neotropical Lepidoptera. Proceedings of the Royal Society B 263:1351-1357. [link]


Adler, G.H., Austin, C.C. and R. Dudley. (1995). Dispersal and speciation of skinks among archipelagos in the tropical Pacific Ocean. Evolutionary Ecology 9:529-541. [link]

Chai, P. and R. Dudley. (1995). Limits to vertebrate locomotor energetics suggested by hummingbirds hovering in heliox. Nature 377:722-725. [link]

Dudley, R. (1995a). Aerodynamics, energetics and reproductive constraints of migratory flight in insects. In: Insect Migration: Tracking Resources Through Space and Time (ed. V.A. Drake and A.G. Gatehouse), pp. 303-319. Cambridge: Cambridge University Press.

Dudley, R. (1995b). Extraordinary flight performance of orchid bees (Apidae: Euglossini) hovering in heliox (80% He/20% O2). Journal of Experimental Biology 198:1065-1070. [link]

Graham, J.B., Dudley, R., Aguilar, N. and C. Gans. (1995). Implications of the late Palaeozoic oxygen pulse for physiology and evolution. Nature 375:117-120. [link]

Larimer, J.L. and R. Dudley. (1995). Accelerational implications of hummingbird display dives. The Auk 112:1064-1066. [link]


Dudley, R. (1994). Aerodynamics of insect dispersal and the constraint of body size. In: Proceedings of the 13th International Biometeorology Congress (ed. A.R. Maarouf, N.N. Barthakur and W.O. Haufe), Part 2 (Vol. 3), pp. 1035-1041. Downsview: Environment Canada.

Dudley, R. and R.B. Srygley. (1994). Flight physiology of Neotropical butterflies: allometry of airspeeds during natural free flight. Journal of Experimental Biology 191:125-139. [link]

Dudley, R. and G.J. Vermeij. (1994). Energetic constraints of folivory: leaf fractionation by frugivorous bats. Functional Ecology 8:668. [link]

Adler, G.H. and R. Dudley. (1994). Butterfly biogeography and endemism on tropical Pacific islands. Biological Journal of the Linnean Society 51:151-162. [link]

Larimer, J.L. and R. Dudley. (1994). Centrifugal force and blood pressure elevation in the wings of flying hummingbirds. Journal of theoretical Biology 168:233-236. [link]


Rand, A.S. and R. Dudley. (1993). Frogs in helium: the anuran vocal sac is not a cavity resonator. Physiological Zoology 66:793-806. [link]

Srygley, R.B. and R. Dudley. (1993). Correlations of the position of center of body mass with butterfly escape tactics. Journal of Experimental Biology 174:155-166. [link]


Dudley, R. (1992). Aerodynamics of flight. In: Biomechanics (Structures & Systems): A Practical Approach (ed. A.A. Biewener), pp. 97-121. Oxford: Oxford University Press.

Dudley, R. and A.S. Rand. (1992). Underwater sound production in a Neotropical anuran, Physalaemus pustulosus (Leptodactylidae). Bioacoustics 4:211-216. [link]

Dudley, R. and G.J. Vermeij. (1992). Do the power requirements of flapping flight constrain folivory in flying animals? Functional Ecology 6:101-104. [link]

Mulkey, S.S., A.P. Smith, S.J. Wright, J.L. Machado and R. Dudley. (1992). Contrasting leaf phenotypes control seasonal variation in water loss in a tropical forest shrub. Proceedings of the National Academy of Sciences USA 89:9084-9088. [link]


Dudley, R. (1991a). Biomechanics of flight in Neotropical butterflies: aerodynamics and mechanical power requirements. Journal of Experimental Biology 159:335-357. [link]

Dudley, R. (1991b). Comparative biomechanics and the evolutionary diversification of flying insect morphology. In: The Unity of Evolutionary Biology (ed. E.C. Dudley), pp. 503-514. Portland: Dioscorides Press.

Dudley, R. (1991c). Thermoregulation in unpalatable danaine butterflies. Functional Ecology 5:503-506. [link]

Dudley, R. and C. Gans. (1991). A critique of symmorphosis and optimality models in physiology. Physiological Zoology 64:627-637. [link]

Dudley, R. and A.S. Rand. (1991). Vocal sac inflation and sound production in the túngara frog, Physalaemus pustulosus (Leptodactylidae). Copeia 1991:460-470. [link]

Dudley, R., King, V.A. and R.J. Wassersug. (1991). The implications of shape and metamorphosis for drag forces on a generalized pond tadpole (Rana catesbeiana). Copeia 1991:252-257. [link]

Stern, D.L. and R. Dudley. (1991). Wing buzzing by territorial male orchid bees, Eulaema meriana (Hymenoptera: Apidae). Journal of the Kansas Entomological Society 64:88-94. [link]


DeVries, P.J. and R. Dudley. (1990). Morphometrics, airspeed, thermoregulation and lipid reserves of migrating Urania fulgens (Uraniidae) moths in natural free flight. Physiological Zoology 63:235-251. [link]

Dudley, R. (1990a). Biomechanics of flight in Neotropical butterflies: morphometrics and kinematics. Journal of Experimental Biology 150:37-53. [link]

Dudley, R. (1990b). Thanatosis in the Neotropical butterfly Caligo illioneus (Nymphalidae: Brassolinae). Journal of Research on the Lepidoptera 28:125-126.

Dudley, R. and P.J. DeVries. (1990a). Flight physiology of migrating Urania fulgens (Uraniidae) moths: kinematics and aerodynamics of natural free flight. Journal of Comparative Physiology A 167:145-154. [link]

Dudley, R. and P.J. DeVries. (1990b). Tropical rain forest structure and the geographical distribution of gliding vertebrates. Biotropica 22:432-434. [link]

Dudley, R. and C.P. Ellington. (1990a). Mechanics of forward flight in bumblebees. I. Kinematics and morphology. Journal of Experimental Biology 148:19-52. [link]

Dudley, R. and C.P. Ellington. (1990b). Mechanics of forward flight in bumblebees. II. Quasi-steady lift and mechanical power requirements. Journal of Experimental Biology 148:53-88. [link]

Dudley, R. and K. Milton. (1990). Parasite deterrence and the energetic costs of slapping in howler monkeys, Alouatta palliata. Journal of Mammalogy 71:463-465. [link]

Dudley, R. (1985). Fluid-dynamic drag of limpet shells. Veliger 28:6-13.

Dudley, R. (1980). Crab-crushing of periwinkle shells, Littorina littorea, from two geographical provinces. Nautilus 94:108-111.